Many of these combining forms may be used as either prefixes or suffixes. Examples are presented to show current usage.
Prefixes | Derived From: | Meaning | Example |
a-, ab- | Latin | off, from, down, away | abduct, avert |
a-, an- | Greek | not, without, less | abiotic, anaerobic |
actin- | G. aktis | a ray, beam, spoke | actinomycete |
ad- | Latin | to, attached to, | adsorption |
aer- | Greek | air | aerobic |
amphi- | Greek | both, about, around | amphibian |
ana- | Latin | away, through, again | analysis |
andro- | Greek | man, male | androgens |
angio- | Greek | a vessel, closed container | angiospermae |
anthropo- | Greek | referring to man | anthropology |
ant-, anti- | Greek | against, away, opposite | antibiosis |
ante- | Latin | before | anteroom |
ap-, aph-, apo- | Latin | from, off, separate | apogee |
aqua- | Latin | water | aquatic |
arche-, archeo- | Greek | ancient, primitive | archeology |
arthri-, arthro- | G. arthron | joint, jointed | arthritis |
asco- | G. askos | bag, sack, bladder | ascospore |
aureo- | L. aureus | gold colored | aureomycin |
auto- | G. autos | self | autoimmune |
bi- | Latin | two, twice, double | bipolar, binocular |
bio-, bios- | Greek | related to life | biology, biocidal |
blasto- | G. blastos | an embryonic layer or cell | blastomere |
brachy- | Greek | short | brachycephalic |
brad-, brady- | Greek | slow, slowness | bradycardia |
bry-, bryo- | G. bryon | moss, mossy | bryophyte |
calic-, calix- | Latin | cuplike | calyx |
cani-, canis- | Latin | dog | canine |
cardia- | G. kardia | heart | cardiac |
carn- | L. carnis | flesh | carnivore |
carp- | L. carpalis | wrist, bones | carpel |
cata- | Greek | decomposition, degradation | catabolism |
cell- | L. cella | small room | cellular |
cephal- | Latin | head | cephalic |
chloro- | G. chloros | green, containing chloride | chlorophyll |
chroma-, chromo- | Greek | colored | chromosome |
chron-, chrono- | G. chronos | time | chronometer |
circum- | Latin | around, near, about | circumnavigate |
coel- | G. koilos | hollow cavity, belly | coelom |
col-, com-, con- | Latin | with, together | combine, collide |
contra- | Latin | against | contradict |
crypto- | G. kryptos | hidden | cryptogamic |
cyano- | G. kyanos | dark blue, blue-green | cyanobacteria |
cyst- | G. kystis | bladder | cystitis |
cyt-,cyte-,cyto- | G. kytos | cell, a hollow vessel | cytology |
de- | Latin | undoing, removal of, from | dehydration |
den-, dent- | L. dens | tooth | dentition |
dendro- | Greek | tree | dendrochronology |
derm-, derma- | Greek | skin, hide | dermatitis |
deut-, deutero- | Greek | second, secondary | deuterium |
di- | Greek | double, twice, two | disaccharide |
dia- | Greek | through, across | diameter |
diplo- | Greek | twofold, double | diploid |
dis- | Latin | apart, away | dissolve |
dorm- | Latin | to sleep | dormant, dormitory |
drom-, drome- | Greek | a running, racing | dromendary |
e-, ec- | Latin | out, out of | efferent |
eco- | G. oikos | house, environment | ecology |
ecto- | G. ektos | outside | ectoderm |
en-, endo- | G. endon | within, internal | endoskeleton |
entero- | G. enteron | intestine | enterocolitis |
entomo- | G. entoma | insect | entomology |
eo-, eos- | Greek | the dawn | Eocene, Eohippus |
epi- | Greek | upon, above, top | epidermis |
erythro- | Greek | red | erythrocyte |
eu- | Greek | proper, true, good | eucaryotic |
ex- | Latin | out, from | excise |
exo- | Greek | outer, external | exoskeleton |
extra- | L. exter | outside of, beyond | extracellular |
flagell- | L. flagrum | whip, whiplike | flagellum |
fuc-, fuco- | G. phyktos | seaweed, algae, lichen | fucoxanthin |
gamo- | G. gamos | sexual union | gamogenesis |
gastero-,gastro- | G. gaster | stomach, belly | gastroenteritis |
geno- | L.gene | origin, development | genotype |
ge-, geo- | Greek | earth | geology |
glu-, glyco- | Greek | sweet, sugar | glucose, glycogen |
gon-,goni-,gono- | Greek | reproductive, sexual | gonorrhea |
gymn-, gymno- | G. gymnos | naked, bare | gymnosperm |
gyn-,gyne-,gyno- | Greek | woman, female | gynecology |
halo- | G. hals | salt | halophile |
haplo- | G. haploos | single | haploid |
heme-, hemo- | G. haimo | blood | hemotologist |
hemi- | Greek | half | hemisphere |
hepta- | Greek | seven | heptane |
herb- | L. herba | pertaining to plants | herbicide |
hetero- | Greek | different, other, unlike | heterozygous |
hex-, hexa- | Greek | six | hexagonal |
hipp-, hippo- | G. hippos | horse | hippodrome |
histo- | G. histos | tissue | histology |
holo- | G. holos | whole, entire | holoblastic |
homeo, homo- | Greek | same, similar, like | homogeneous |
hyal-, hyalo- | G. hyalos | glassy, transparent | hyaloid |
hydr-, hydro- | Greek | pertaining to water | hydrolysis |
hyper- | Greek | above, more, over | hyperactive |
hypo- | Greek | below, less, under | hypodermic |
ichthy-,ichthyo- | Greek | referring to fish | ichthyology |
inter- | Latin | between | intercellular |
intra- | Latin | within, inside | intracellular |
intro- | Latin | inward, within | introvert |
iso- | Greek | equal, same | isotonic |
kine- | Greek | movement, moving | kinetics |
leuc-, leuk- | Greek | white | leucocyte |
lycan- | G. lykos | wolf | lycanthropy |
macro- | Greek | large, big, long | macromolecule |
man-, manu- | Latin | hand | manual |
mastig- | G. mastigos | whip | mastigophora |
meg-, mega- | Greek | great, large | megabyte |
melan-,melano- | Greek | black, dark | melanin |
mero- | G. merus | part, piece | meroblast |
mes-, meso- | G. mesos | middle, in between | mesoderm |
met-, meta- | Greek | later, following, changed in position or form | metamorphosis |
micro- | G. mikros | small | microbiology |
milli- | Latin | a thousandth part | millimeter |
mio- | G. meion | less, smaller | Miocene |
mito- | G. mitos | thread | mitosis |
mon-, mono- | Greek | one, single | monocular |
morph- | Greek | shape, form | morphology |
mor-, mort- | Latin | die, death, | mortality |
muc-, muco- | Latin | consisting of many units | multicellular |
mus- | Latin | mouse, as one running | muscle |
myco-, mykos- | Greek | fungus, mushroom | mycology |
myo- | G. mys | muscle | myoglobin |
myxo- | Greek | slime, mucus | myxomycetes |
nemato- | Greek | thread, threadlike | nematode |
neuro- | Greek | name | nomenclature |
ob- | Latin | against | obtuse |
octa- | Greek | eight | octopus |
olig-, oligo- | Greek | few, small, less | oligarchy |
omni- | Latin | all, everywhere | omnipotent |
oo- | Greek | pertaining to an egg | oocyte |
ophthalmo- | Greek | referring to the eye | ophthalmologist |
opisth-,opistho- | Greek | behind, backwards, back | Opisthobranchia |
orni-, ornitho- | Greek | bird | ornithology |
orth-, ortho- | Greek | straight | orthodontist |
osteo- | Greek | bone | osteocyte |
oto- | Greek | referring to the ear | otology |
ova-,ovi-,ovul- | Latin | egg | ovary, oviduct |
paleo- | Greek | old, ancient | paleontology |
para- | Greek | beside, near, beyond | parasitism |
path-, patho- | Greek | disease, suffer | pathogenic |
ped-, pedi- | Latin | foot | pedicure |
penna-, pinna- | Latin | feather, feathery | pinnate |
pent-, penta- | Greek | five | pentagon |
per- | Latin | through | pervade, peruse |
peri- | Greek | around, surrounding | perimeter |
pher- | Greek | bearing, carrying, support | pheromone |
phil- philo- | Greek | loving, attracted to | philanthropy |
phob- | Greek | fear, fearing | phobic |
photo- | Greek | pertaining to light | photosynthesis |
phyco- | Greek | seaweed, algae | phycology |
phylo- | Greek | tribe, race, related group | phylogeny |
phyto- | Greek | pertaining to plants | phytohormone |
plasm-, plasma- | Greek | formative substance | plasmablasts |
plati-, platy- | Greek | flat | platypus |
pleio- pleo- | Greek | more, many | pleiomorphic |
pod-,poda-,podi- | Greek | foot | podiatrist |
poly- | Greek | many | polyhedron |
post- | Latin | after | postnatal |
pre- | Latin | before | prenatal |
preter- | Latin | beyond | preterhuman |
prim- | Latin | first | primary |
pro- | Greek | before, on behalf of | proboscis |
pro- | Latin | forward | progressive |
proto- | Greek | first, primary | protozoa |
pseudo- | Greek | false | pseudopod |
psilo- | Greek | bare, mere | psilopsida |
pteri-, ptero- | Greek | fern, feather | pteridophyte |
quadr-, quadri- | Latin | four | quadruped |
radi- | Latin | ray, spoke of wheel | radial |
re- | Latin | back, again | repeat |
retro- | Latin | backward | retroactive |
rhiz-, rhizo- | Greek | pertaining to roots | rhizoids |
rhod-, rhodo- | Greek | a rose, red | rhodopsin |
rota- | Latin | wheel | rotate |
sapr-, sapro- | Greek | rotten, putrid, dead | saprobe |
sarc-, sarco- | Greek | flesh, fleshy | sarcoma |
schiz-, schizo- | Greek | split, splitting | schizocoel |
se- | Latin | apart | secede |
semi- | Latin | half | semicircle |
soma-, somato- | Greek | body | somatic |
sperma-,spermato- | Greek | seed | spermatozoa |
sporo- | Greek | spore | sporophyte |
staphylo- | Greek | bunch of grapes | staphylococcus |
stoma- | Greek | mouth | stomate |
strepto- | Greek | twisted, string of | streptococcus |
sub- | Latin | below, under, smaller | subapical |
supra-, super- | Latin | above, over | supernova |
sym-, syn- | Greek | together, with | synthesis |
taxi-, taxo- | Greek | to make order, arrangement | taxonomy |
tel-,tele-,telo- | Greek | distant, end | telophase |
terra-, terre- | Latin | land, earth | terrestrial |
tetra- | Greek | four | tetrapod |
therm-, thermo- | Greek | heat | thermometer |
thigmo- | Greek | touch | thigmotaxis |
trans- | Latin | across, through, over | transfer |
tri- | Latin | three | triangle |
tricho- | Greek | hair | trichocyst |
triplo- | Latin | triple | triploid |
troche-, trocho- | Greek | wheel, hoop | trochophore |
tropho- | Greek | nourishment | trophoblast |
ultra- | Latin | beyond, exceedingly | ultraconservative |
uni- | Latin | consisting of one | unicellular |
vice- | Latin | in place of | vice-president |
vid-, vis- | Latin | see | vision |
xen-, xeno- | Greek | dry, desert | xerophyte |
zoo- | Greek | animal, life | zoology |
zyg-, zygo- | Greek | to join together | zygote |
Suffixes | Derived From: | Meaning | Example |
-biosis | Greek | mode of living, way of life | symbiosis |
-blast | Greek | formative, embryonic | mesoblast |
-chaeta-, -chete | Greek | a bristle | Polychaeta |
-chrome | Greek | color | mercurochrome |
-cidal, -cide | Latin | killer, a killing | insecticide |
-cocci, -coccus | Greek | round, seed, kernel | Streptococcus |
-cyst | Greek | pouch, sac | trichocyst |
-dactyl | Greek | finger | pentadactyl |
-derm, -dermis | Greek | skin, layer | epidermis |
-elle, -ule, -la, -le, -let, -ole | Latin | small, diminutive endings | globule, piglet |
-emia | Greek | blood disease | anemia |
-fer | Latin | bearer, producer, carry | conifer, transfer |
-gamous, -gamy | Greek | marriage, sexual fusion | polygamy |
-gen, -geny | Greek | origin, production | progeny, hydrogen |
-genesis | Latin | origin, development of | embryogenesis |
-gony | Latin | something produced | cosmogeny |
-graph | Greek | drawing, writing | chromatograph |
-hedral, -hedron | Greek | side | polyhedral |
-hydrate | Greek | compound formed by union of water with other substance | carbohydrate |
-ism | Greek | act, practice or result of | terrorism |
-ite | Latin | a division or part | somite |
-itis | Greek | inflammation or infection | appendicitis |
-jugal, -jugate | Latin | to yoke, join together | conjugate |
-logy | G. logos | science or study of | biology |
-lysis, -lytic | Greek | loosening, separation, splitting into smaller units | photolysis |
-mer, -merous | G. meros | a part, piece | polymer |
-meter | G. metron | a measurement | diameter |
-morph | Greek | form | endomorph |
-mycin | Greek | derived from a fungus | aureomycin |
-nomy | Greek | systematized knowledge of | astronomy |
-oma | Greek | tumorous | carcinoma |
-osis, -otic | Greek | abnormal condition, disease | neurosis |
-phage | Greek | eater | bacteriophage |
-phase | Greek | a stage or condition | metaphase |
-phil, -phile | Greek | fear, fearing | hydrophobia |
-phor, -phore | Greek | bearing, carrying, supporting | sporangiophore |
-phyll | Greek | leaf | chorophyll |
-phyta, -phyte | Greek | plant | epiphyte |
-plasm | Greek | formative substance | cytoplasm |
-plast | Greek | organized particle, granule | choroplast |
-pod, -poda | Greek | foot | arthropod |
-some | Greek | body | chromosome |
-stasis | Greek | a stationary position | homeostasis |
-stat, -static | Greek | stationary, still | hemostat |
-stomy | Greek | opening into | colostomy |
-therm | Greek | heat | homeotherm |
-thes, -thesis | Greek | arrangement, in order | hypothesis |
-tom, -tomy | Greek | dividing, surgery | lobotomy |
-trope, -tropic | Greek | turning | phototropic |
-vor, -vore | L. vorare | feeding | carnivore |
-zoa,-zoan,-zoic | Greek | animal, life | protozoa |