Nicolaidou Artemis


ďel. 727-4359, Fax: 727-4359


University education - Courses taught - Research interests - Recent Research Projects

Ph D Theses supervised - Publications

"Thalis" Images




University education

B.Sc. Natural Science, University of Athens, 1972
M.Sc. Marine Biology, University of Wales, Bangor, UK, 1974
Ph.D. Marine Biology, University of Wales, Bangor, UK, 1977

Courses taught


  • Biological Oceanography
  • Zoology


  • Benthic Ecology
  • Estimation of biodiversity

Research Interests

  • Benthic Ecology
  • Ecology of coastal lagoons
  • Impact of pollution on benthic communities and populations
  • Polychaete taxonomy







Recent Research Projects

  • 2000. Ecological quality of the water of Maliakos Gulf. (Ministry of the Environment).
  • 2002. Import of alien species through shipping in greek ports and effects on native ecosystems (General Secretariat of Research and Technology).
  • 2004. Environmental stress and benthic communities of brackish water lagoons: effects on biodiversity and life strategies (Ministry of Education).
  • 2005. Management and sustainable development of protected transitional waters (EU, INTEREG).
  • 2005. The role of the microbial community in the decomposition of terrestrial organic matter in transitional waters (General Secretariat of Research and Technology).
  • 2006. Geographical isolation of greek lagoons and genetic diversity of their populations (University of Athens).







Ph D Theses supervised

  • Simboura, N. 1995 Marine Benthic Polychaetes of Greece: Taxonomy, Ecology Zoogeography.
  • Pancucci-Papadopoulou, A. 1996.Ecology and distribution of the echinoderms in Greece. Dwarfism in the sea-urchin Paracentrotus lividus (Lamark) as an ecological adaptation.
  • Panagopoulos, D. 1996. The effects of solid wastes on benthic communities.
  • Reizopoulou, S. 1997. The zoobenthos of brackish water lagoons. Variability in relation to environmental factors.
  • Kormas, K.1998. Description and dynamics of the ecological components of the ecosystem of Maliakos Bay.
  • Vardala-Theodorou, E. 1998. Study of the benthic mollusks, living and fossil, from Vouliagmeni Lake, Perahora.
  • Tsagari, A. 2001. Effect of pollution on the physiology, morphology and biochemistry of Mytilus galloprovincialis.
  • Kambouroglou, V. 2009. Import of alien species through shipping in greek ports and effects on native ecosystems.






























































































1974 MSc Thesis Influence of sediment properties and wave action on tube building and distribution of Lanice conchilega, University of Wales. 56 pp

1977 Ph.D. Thesis Variability in Pectinaria dominated benthic communities, University of Wales 172 pp



1. Simboura N. and A. Nicolaidou, 2001. The Polychaetes (Annelida, Polychaeta) of Greece: Checklist, distribution and Ecological Characteristics. Monographs on Marine Sciences 4. 115 pp. National Centre for Marine Research.

2. Nicolaidou A., S. Reizopoulou, D. Koutsoubas S., Orfanidis & Th. Kevrekidis 2005. Lagoons pp.3-11, VI.1. In: State of the Hellenic Marine Environment. E. Papathanassiou & A. Zenetos (eds), Inst. Oceanoraphy, HCMR, 2005, 358 pp.

3. Nicolaidou A., S. Reizopoulou, D. Koutsoubas S., Orfanidis & Th.Kevrekidis, 2005. Biological components of Greek lagoonal ecosystems: a first synthesis. Mediterranean Marine Science 6: 31-50. PDF



1. Rees, E.I.S., A. Nicolaidou and P.Laskaridou, 1977. The effects of storms on the dynamics of shallow water benthic associations. In Keegan, B.F., O'Ceidigh and P.J.Boaden (Eds). Biology of Benthic Organisms. Pergamon Press, p.465-477.

2. Nicolaidou, A., 1983. Life history and productivity of Pectinaria koreni Malmgren (Polychaeta). Estuar. coast. Shelf Sci. 17: 31-43. PDF

3. Nicolaidou, A., M. Moraitou-Apostolopoulou and L. Ignatiades, 1983. A survey of estuarine benthic, zooplanktonic and phytoplanktonic communities of Amvrakikos Gulf, Ionian Sea. Mar. Ecol. 4: 197-209.

4. Taraschewski, H. and A. Nicolaidou, 1987. Heterophyes species in Greece: record of H. heterophyes, H.aequalis and H. dispar from the first intermediate host. J. Helminth. 61: 28-32.

5. Karakiri, M. and A. Nicolaidou, 1987. Population studies on the Amphipoda of Mazoma lagoon, Greece. Helgolander meeresunters 41: 453-464.

6. Karakiri, M. and A. Nicolaidou, 1988. On a collection of Amphipoda from the Northern Sporades, Aegean Sea. Crustaceana, Sup. 13: 107-114

7. Nicolaidou, A. 1988. Notes on the behaviour of Pectinaria koreni Malmgren. J.mar. biol. Assoc., U.K., 68: 55-59.

8. Nicolaidou, A. and M. Kostaki- Apostolopoulou, 1988. Growth of Abra ovata in a brackish-water lagoon. Vie marine, 9: 7-10.

9. Nicolaidou, A., F. Bourgoutzani, A. Zenetos, O. Guelorget and J.-MP. Perthuisot, 1988. Distribution of Molluscs and Polychaetes in coastal lagoons in Greece. Estuar. coast. Shelf Sci., 26: 337-350. PDF

10. Nicolaidou, A. and M. Karakiri, 1989. The distribution of Amphipoda in a brackish water lagoon in Greece. Mar. Ecol., 10: 131-139.

11. Nicolaidou, A., A.M.Pancucci and A. Zenetos, 1989. The impact of dumping coarse metalliferous waste on the benthos in Evoikos Gulf, Greece. Mar.Pollut. Bull., 20: 28-33. PDF

12. ═icolaidou, A. and C-N. Papadopoulou, 1989. Factors affecting the distribution and diversity of polychaetes in Amvrakikos Bay, Greece. Mar. Ecol. 11: 193-204.

13. Nicolaidou, A. and J.A. Nott, 1989. Heavy metal pollution induced by a ferro-nickel smelting plant in Greece. Sci. Total Envir. 84: 113-117. PDF

14. Nott, J.A. and A. Nicolaidou, 1989. The cytology of heavy metal accumulations in the digestive glands of three marine gastropods. Proc. R. Soc. Lond. B. 237: 347-362.

15. Panagopoulos, D. and A. Nicolaidou, 1989. A population of Desdemona ornata Banse 1957 (Polychaeta, Sabellidae) settled in a fully marine habitat of the Mediterranean. Oebalia, 15: 35-39.

16. Nott, J.A and A. Nicolaidou, 1989. Metals in gastropods: metabolism and bioreduction. Mar. environ. Res., 28: 201-205.

17. Nott, J.A. and A. Nicolaidou, 1990. Transfer of metal detoxification along marine food chains. J.mar. biol. Assoc., U.K., 70: 905-912.

18. Nicolaidou, A. and J.A. Nott, 1990. Mediterranean pollution from a ferro-nickel smelter: Differential uptake of metals by some gastropods. Mar. Pollut. Bull. 21: 137-143.

19. Zenetos, A., F. Bei and A. Nicolaidou, 1991. Erratic occurrence of benthic fauna in a shallow Mediterranean area: an indirect effect of manmade disturbance. Mar. Pollut. Bull. 22: 618-622. PDF

20. Nott, J.A. and A. Nicolaidou, 1993. Bioreduction of zinc and manganese along a molluscan food chain. Comp. Biochem. Physiol. 104a: 235-238. PDF

21. Nicolaidou, A., A. Zenetos, M.-A. Pancucci and N.Simboura, 1993. Comparing ecological effects of two different types of pollution using multivariate techniques. Mar. Ecol., 14: 113-128.

22. Catsiki, A.B., F. Bei and A. Nicolaidou, 1994. Size dependent metal concentrations in two marine gastropods. Neth. J. Aquat. Ecol.,28: 157-165.

23. Nott, J.A. and A. Nicolaidou, 1994. Variable transfer of detoxified metals from snails to hermit crabs in marine food chains. Mar. Biol. 120: 369-377.

24. Simboura, N., A.Zenetos, M.Thessalou-Legaki, A.Pancucci-Papadopoulou and A.Nicolaidou, 1995. Benthic communities of the infralittoral in the N. Sporades (Aegean Sea). A variety of biotopes encountered and analysed. Mar. Ecol. 16:283-306.

25. Reizopoulou, S., M. Thessalou-Legaki and A. Nicolaidou, 1996. Assessment of disturbance in Mediterranean lagoons: an evaluation of methods. Mar. Biol. 125: 189-197. PDF

26. Nott, J.A. and A.Nicolaidou, 1996. Kinetics of metals in molluscan faecal pellets and mineralised granules, incubated in marine sediments. J.exper.mar. Biol. Ecol.197:203-218. PDF

27. Kormas, K.A., M. Thessalou-Legaki and A.Nicolaidou, 1997. Is water column productivity in an Aegean embayment reflected to its benthic assemblages? In: Hawkins, L.E., S. Hutchinson, A.C. Jensen, M. Sheader and J.A. Williams (Eds.) The response of marine organisms to their environment. Univ. of Southampton, pp.143-151.

28. Nicolaidou, A. and J..A. Nott, 1998. Metals in sediment, sea grass and gastropods near a nickel smelter in Greece: Possible interactions. Mar. Pollut. Bull. 35: 360-365. PDF

29. Kormas, K.A., K. Kapiris, M. Thessalou-Legaki and A.Nicolaidou, 1998. Quantitative relationships between phytoplankton, bacteria and protists in an Aegean semi-enclosed embayment (Maliakos Gulf, Greece). Aquat. Microb. Ecol 15: 255-264.

30. Gibbs, P.E., J.A. Nott, A. Nicolaidou and M.J. Bebianno, 1998. The composition of phosphate granules in the digestive glands of marine prosobranch gastropods: variation in relation to taxonomy. J. Moll. Stud. 64: 423-433. IF:0.968

31. Nicolaidou, A. and J.A. Nott 1999. The role of the marine gastropod Cerithium vulgatum in the biogeochemical cycling of metals. In: J.S. Gray et al (eds) Biogeochemical Cycling and Sediment Ecology. Kluwer Academic Publishers, Netherlands. pp:137-146.

32. Pancucci-Papadopoulou, A., N.Simboura, A.Zenetos, M.Thessalou-Legaki and A. Nicolaidou 1999. Benthic invertebrate communities of NW Rodos (Rhodes) Island (SE Aegean Sea) as related to hydrological regime and geographic location. Israel J. Zool. 45: 371-393.

33. Simboura, N., ┴. Nicolaidou and M. Thessalou-Legaki 2000. Polychaete communities of Greece: An ecological overview. Mar. Ecol.21: 129-144. PDF

34. Makra, A. and A. Nicolaidou, 2000. Benthic communities of the inner Argolikos Bay. Belgian J. Zool. 130 (Sup): 63-69.

35. Kormas, K.A., A. Nicolaidou and S. Reizopoulou, 2001. Temporal variations of nutrients, chlorophyll a and particulate organic matter in three coastal lagoons of Amvrakikos Gulf (Ionian Sea, Greece). Mar. Ecol. 22: 201-213. PDF

36. Makra, A., M. Thessalou-Legaki, J. Costelloe, A. Nicolaidou and B.F. Keegan, 2001. Mapping the pollution gradient of the Saronikos Gulf Benthos prior to the operation of the Athens sewage treatment plant, Greece. Mar. Pollut. Bull. 42: 1417-1419. PDF

37. Kormas, K., V. Garametsi and A. Nicolaidou, 2002. Size fractionated phytoplankton chlorophyll in an Easter Mediterranean coastal system (Maliakos Gulf, Greece). Helgol. Mar. Res. 56:125-133.

38. Brown, C.J., R.A. Eaton, S.M. Cragg, P. Goulletquer, A. Nicolaidou, M.J. Bebianno, J. Icely, G. Daniel, T. Nilsson, A.J. Pitman and G.S. Sawyer, 2003. Assessment of effects of chromated copper arsenate (CCA)-treated timber on nontarget epibiota by investigation of fouling community development at seven European sites. Arch. Environ. Contam. Toxicol. 45: 37-47

39. Nicolaidou, A. 2003. Observations on the re-establishment and tube construction by adults of the polychaete Lanice conchilega. J. Mar. Biol. Ass. U.K. 83:1223-1224.

40. Kormas, K., A. Nicolaidou and M.Thessalou- Legaki 2003. Variability of environmental factors of an eastern Mediterranean Sea river-influenced coastal system. Medit. Mar. Sci. 4:67-77. PDF

41. Antoniadou, C., A. Nicolaidou and C. Chintiroglou. 2004. Polychaetes associated with sciaphilic alga community in the northern Aegean Sea: spatial and temporal variability. Helg. Mar. Res. 58:168-182. PDF

42. Reizopoulou, S. & A. Nicolaidou, 2004. Benthic diversity of coastal brackish-water lagoons in Western Greece. Aquatic Conserv: Mar. Freshw. Ecosyst. 14: S93-S102. PDF

43. Nicolaidou, A., K. Petrou, K.A.Kormas and S. Reizopoulou. 2006. Inter-annual variability of soft bottom macrofaunal communities in two Ionian Sea lagoons. Hydrobiologia 55:89-98. PDF

44. Akoumianaki, I., S. Papaspyrou & A. Nicolaidou, 2006. Dynamics of macrofauna body size in a delta-front environment. Mar. Ecol. Prog. Ser. 321:55-66. PDF

45. Reizopoulou, S. and A. Nicolaidou, 2007. Index of size distribution (ISD): a method of quality assessment for coastal lagoons. Hydrobiologia 577: 141-149. PDF

46. Akoumianaki, I. & A. Nicolaidou, 2007. The spatial variability and dynamics of macrobenthos in a Mediterranean delta front area: role of physical processes. J. Sea Res. PDF

47. Katsanevakis, S. G. Verriopoulos, A. Nicolaidou & M. Thessalou-Legaki, 2007. Effect of marine litter on the benthic megafauna of coastal soft bottoms: a manipulative field experiment. Mar. Pollut. Bull. 54:771-778. PDF

48. Vardala-Theodorou, E., A. Nicolaidou, 2007. On the recent and fossil malacofauna of "Vouoliagmeni Lake", Perachora (Korinthiakos Gulf, Greece). Boll. Malacol. 43: 62-70.

49. Nicolaidou, A. 2007. Lack of temporal variability in the benthos of a coastal brackish water lagoon in greece. Medit. Mar. Sci. 8:5-17. PDF

50. Faulwetter, S., Gotsis, P., Reizopoulou, S., Orfanidis, S., Kevrekidis, T., Nicolaidou, A., Simboura, N., Malea, P. , Dounas, C. , Mogias, A. , Valavanis, V. , Arvanitidis, C. 2008 ElNet: The Greek biodiversity transitional waters information system. An exemplar for the development of distributed information networks in Europe Aquatic Conservation: Marine and Freshwater Ecosystems 18: S135-S142. PDF

51. Sangiorgio, F, Basset, L., M. Pinna, A., Sabetta, Abbiati, M. M. Ponti, M. Minocci, S. Orfanidis, A. Nicolaidou, S. Moncheva, A. Trajanova, L. Georgescu, S. Dragan, S. Beqiraj, D. Koutsoubas, A. Evagelopoulos, & S. Reizopoulou. 2008. Environmental factors affecting fragmites australis litter decomposition in Mediterranean and Black Sea transitional waters. Aquatic Conserv: Mar. Freshw. Ecosyst. 18: S16-S26. PDF

52. Tsangaris, C., E. Papathanassiou & A. Nicolaidou, 2008. Biochemical biomarkers and overall health status of mussels Mytilus galloprovincialis exposed to nickel and chromium. Chemistry and Ecology 24:315-327. PDF

53. Basset, A., Sabetta, L., F. Sangiorgio, M. Pinna, D. Migoni, F. Fanizzi, E. Barbone, N. Galuppo, S. Fonda Umani, S. Reizopoulou, A. Nicolaidou, C. Arvanitidis, S. Moncheva, A. Trajanova, L. Georgescu & S. Beqiraj. 2008. Biodiversity conservation in Mediterranean and Black Sea lagoons: a trait-oriented approach to benthic invertebrate guilds. Aquatic Conserv: Mar. Freshw. Ecosyst. 18: S4-S15

54. Somerfield, P.J. Arvanitidis, C. , Faulwetter, S. , Chatzigeorgiou, G. , Vasileiadou, A. , Amouroux, J.M. , Anisimova, N. , Cochrane, S.J. , Craeymeersch, J. , Dahle, S. , Denisenko, S. , Dounas, K. , Duineveld, G. ,Gremare, A. , Heip, C.H.R. , Herrmann, M., Karakassis, I. , Kedra, M., Kendall, M.A. , Kingston, P. , Kotwicki, L. , Labrune, C. , Laudien, J. , Nevrova, H. , Nicolaidou, A. , Occhipinti-Ambrogi, A. , Palerud, R. , Petrov, A. , Rachor, E. , Revkov, N. , Rumohr, H. , Sard?, R. , Janas, U. , Vanden Berghe, E. , W?odarska-Kowalczuk, M. 2009. Assessing evidence for random assembly of marine benthic communities from regional species pools Marine Ecology Progress Series 382, pp. 279-286

55. Karayanni, H., Kormas, K.Ar., Cragg, S., Nicolaidou, A. 2009 Establishment and Succession of an Epibiotic Community on Chromated Copper Arsenate-Treated Wood in Mediterranean Waters Archives of Environmental Contamination and Toxicology, pp. 1-8. PDF



1. Kambouroglou V. & A. Nicolaidou, 2006. A new alien species in Hellenic waters: Pseudonereis anomala (Polychaeta, Nereididae) invades harbors in the Eastern Mediterranean. Aquatic Invasions 1:97-98. PDF

2. Thessalou-Legaki, M., A. Zenetos, V. Kambouroglou, Corsini-Foka M., P. Kouraklis, C. Dounas & A. Nicolaidou, 2006. The establishment of the invasive crab Percnon gibbesi (H. Milne Edwards, 1853) (Crustacea: Decapoda: Grapsidae) in Greek waters. Aquatic Invasions, 1: 133-136.

3. Sangiorgio F, A. Basset, M. Pinna1, L. Sabetta1, M. Abbiati, M. Ponti, M. Minocci, S. Orfanidis, A. Nicolaidou, S. Moncheva6, A. Trayanova, L. Georgescu, S. Dragan, S. Beqiraj, D. Koutsoubas, A. Evagelopoulos, S. Reizopoulou 2007. Ecosystem processes: litter breakdown patterns in Mediterranean and Black Sea transitional waters Transit. Waters Bull. 3(2007), 51-55. PDF

4. Reizopoulou, S & A. Nicolaidou, 2007. The growth of the bivalve Abra segmentum (Recluz, 1843) in two Mediterranean lagoons. Transit. Waters Bull. 4.